The 4.00 mm pitch and 3.50 mm row spacing give it enough room for the rubber wire seals that keep moisture and dust out of the cavity — the housing itself is listed as sealed, meaning the interface between the receptacle and its mating header is protected against splash and spray when properly assembled with the correct seals and contacts. The gray PBT, glass-filled shell handles the -40°C to 120°C operating range common to underhood and industrial cabinet applications.
The housing itself has no latches or locking ramps listed in the spec, so the mating header's locking feature provides the retention.